Bi 321 (2010)                        Early Evolution of Flowering Plants                 Dr. Dutton

 

Angiosperms

•    Their Monophyly is …

•     

 

•    They probably originated during the Jurassic (> 135 mya)

•    What about “groups” within the Angiosperms?

•    The Monocots (i.e., Liliopsida)

•     

 

 

•    The Dicots (i.e., Magnoliopsida), however …

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•    Understanding Figures 12-1 & 12-2

 

Data used to construct these Figures

•    Figure 12-1

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–  

 

•    Figure 12-2

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–  Tricolpate pollen (and derived types)

 

 

–  describes pollen grains with three ____________________, three compound

 

 

____________________ or three ____________________

 

Your Text: Vascular Plant Taxonomy

•    Provides, for each FAMILY covered:

•    Description

•    Floral formula

•    Distribution and Ecology

•    Number of genera and species and major genera

•    Economic plants and products

•    Discussion

•    MY RECOMMENDATION: Use your text as a supplementary source of information and USE THE DIAGRAMS

 

Resolving “Conflict” Between Lecture and Lab

•    Magnoliidae (sensu Cronquist) equals … “Magnoliid complex”, some members of “Basal Families”, and some members of the Tricolpates (Eudicots) (sensu your Text)  

 

 Magnoliidae  - Features

•     Uniaperturate  pollen

•     Apocarpous

•     Gynoecium  and numerous stamens in a centripetal  sequence (i.e., development from the outside toward the inside or from the apex to the base)

•     Petals are free

•     Some with  vesselless  wood, laminar stamens and more than two cotyledons

•     Many have volatile oils

 

Aristolochiaceae  ( Magnoliids )

•    Herbs, shrubs or woody vines

•    Leaves alternate, simple, often cordate, estipulate

•    CA or z 3 rarely 4 C0-0(3) A4-many GI(3-6); fruits capsules

•    8-10 genera, 600 species

•    Mostly tropical, few temperate

•    Our Flora: Asarum

 

 

Nymphaeaceae  (Basal Angiosperms)

•    Aquatic herbs with long-petioled floating leaves arising from stout rhizomes and long-peduncled flowers bearing numerous parts

•    Leaves alternate, cordate or peltate

•    CA4-many C08-many A-many GS(3-many); fruits spongy berries

•    5 genera, 50 species

•    Widespread

•    Many monocotyledonous features; widely cultivated

•    Our Flora: Brasenia, Nuphar

 

Ranunculaceae  (Basal  Tricoplates )

•    Herbs with bisexual flowers of numerous spirally arranged parts

•    Leaves usually alternate, simple or compound, estipulate

•    CA3-many C00-many A-many GS3-many; fruits achenes follicles, berries, rarely capsules

•    Ca 50 genera, 2000 species

•    Cosmopolitan but mainly temperate and boreal

•    Many cultivated and some medicinal species although some very poisonous. Some species of Ranunculus are weedy.

•    Our Flora: Aconitum, Actaea, Anemone, Aquilegia, Caltha, Cimicifuga, Clematis, Coptis, Delphinium, Isopyrum, Myosurus, Ranunculus, Thalictrum, Trautvetteria»

 

Berberidaceae  (Basal  Tricoplates )

•    Herbs or shrubs with biseriate stamens opposite the petals and anthers dehiscing by flaps

•    Leaves alternate, simple or compound, generally estipulate

•    CA6-9, usually 3+3 C06-9, usually 3+3 A6(4-18) GS(2-3); fruits berries

•    13 genera, 650 species

•    Mainly north temperate regions.

•    Note: Poorly defined by Cronquist

•    Our Flora: Achlys, Berberis, Vancouveria

 

Papaveraceae  (Basal  Tricoplates )

•    Herbs with milky or colored sap, bisexual flowers with deciduous sepals and numerous stamens in several whorls

•    Leaves alternate or basal, simple or compound, estipulate

•    CA2-4 C04or6(8, 12, 16, or 0) A-many GS(2-many); fruits capsules

•    25 genera, 200 species

•    Mainly north temperate regions

•    Cultivated and medicinal plants, especially Papaver somniferum (opium)

•    Our Flora: Eschscholtzia, Meconella

 

Fumariaceae  (Basal  Tricoplates )

•    Included in Papaveraceae by Walters et al.

•    Herbs with watery sap, generally dissected leaves and zygomorphic flowers with deciduous sepals, paired petals and a compound ovary

•    Leaves alternate or basal, simple or compound, estipulate

•    CA2 COz2+2 A6 GS(2); fruits capsules or nuts

•    19 genera, 400 species

•    Mainly north temperate regions

•    Some cultivated, several weedy (Corydalis)

•    Our Flora: Corydalis, Dicentra, Fumaria

 

Important Note:

•    I will not be providing additional lecture guides until the 9th week of the term

– the characters discussed here are included in your lab packets

– bring your lab packets to lecture!